This meadowlark occurs in dry fields, upland pastures and grasslands with few or no trees. It delivers its beautiful song from a fence post or high point in its territory during the summer. At this time it eats mostly insects and spiders. The cup-like nest is on the ground and covered with a roughly woven dome of long grasses. It is regularly parasitized by Brown-headed Cowbird eggs. During the winter, the Western Meadowlark feeds on seeds, forms small flocks and roosts at night on the ground.
The four-digit banding code is WEME.